ZUTTA, B.R, D.SIMS and J.AGAMON
California State University of Los Angeles, Los Angeles, CA 90032 USA
Southern Californian vegetation, exposed to strong seasonal weather cycles and dynamic disturbance regimes, undergo wide variations in photosynthetic fluxes and other ecosystem processes. Sorting vegetation by "functional types," based on their resource use patterns and seasonal response, provides one simplification to help understand changing ecosystem fluxes in these systems. For this approach to be broadly applicable, these functional types must be detectable through remote sensing. Our objective was to measure the seasonal change in spectral reflectance of Southern Californian vegetation and to use the seasonal and spectral variability to categorize species into optically distinct types. A field spectrometer was used to measure reflectance at 1-meter intervals along 100 meter transects in a variety of vegetation types, located in the Santa Monica Mountains of Southern California. Several spectral indices were used in principal components analysis to characterize the variability in spectral reflectance in time and space. Initial results of this study indicate limited power to separate functional groups (e.g. associated with contrasting phenology or photosynthetic rates) when spectra from a single date were used. The ability to separate optical types was enhanced when multiple sampling dates were included in the analysis. Further work is focusing on the physiological characteristics of the optically determined functional groups. Development of this approach may lead to better characterization of functional types and an understanding of the impacts of disturbance and global change on ecosystem processes.
INTRODUCTION
Reflectance indices can be used to show contrasting spectral patterns of distinct vegetation types or reflect seasonal changes.
This study proposes a new way to interpret remotely sensed data to understand large scale vegetation patterns and processes. Further development of this approach can be applied for understanding the impacts of disturbance and global change.
Objective
1- Categorize southern Califonian vegetation into functional types through their optical response, detectable with remote sensing.
2 - Examine functional type seasonal changes.
METHODS
Optical sampling (remote sensing) from the ground employs a spectrometer,
held above the canopy, and samples vegetation along a transect or within
a quadrat. Eight 100 meter transects were established at each site to represent
the dominant functional types of the area. These types included annuals,
drought deciduous, evergreen, and winter deciduous vegetation. Each point
was sampled repeatedly through the study period.
Site Locations

Field Spectrometry

Optical View of Vegetation

Figure 2. Reflectance indices can be used to show contrasting
spectral patterns of distinct vegetation types.
Table 1. Examples of common physiological indices derived from "optical"
sampling (see Figure 1).
From Two Dimensions to a Multi-Dimensional Graph



Dominant Southern Californian Vegetation

Species and Functional Type

Seasonal Changes of Vegetation

RESULTS/DISCUSSION
Categorizing functional types through their spectral reflectance
Spectral indices and principle component factors were used to discriminate optical patterns into separate groups (See Table 2 for functional type list) for each month.
We found that many functional types could be distinguished optically from one another at seasonal extremes. After high precipitation in February and high water stress in July, groups seperated more due to changes in canopy structure (see Figure 6 and 10).
Figure 6. The beginning of winter initiated new growth in annuals.
Many evergreens and deciduous vegetation responded similar to each other.
Figure 7. The unusual lack of precipitation in January had stressed
many plants types. Evergreen shrubs (i.e. Malosma laurina) showed
little physical effect.
Figure 8. The return of precipitation initiated a large quantity
of growth. All functional reacted with vegetative growth.


Figure 11. Increased water stress forced many functional types
decline. Winter deciduous vegetation were still photosynthetically active.

Summary of seasonal changes

Figure 14. Seasonal location of functional types were related to changes
in their canopy structure.

CONCLUSIONS
1- Spectral classification of functional types worked best after the high precipitation of early spring and the sever water stress of mid-summer.
2- Seasonal changes in annuals and winter deciduous canopy structures allowed for relatively distinct spectral groups throughout the year. Drought deciduous and evergreen vegetation mixed regularly.
3 - Additional research is needed to determine if spectral classification
of functional types change during episodes of exteme drought or percipitation.
ACKNOWLEDGEMENTS
We would like to thank Cea-Crest (Center for Environmental Analysis) for their continued support of this study, Dafna Kohn and the National Park service for supplying maps, and the students and professors of the California State University, Los Angeles for their continued support.